Il temolo artico, Thymallus arcticus (Pallas, 1776)

Nomi comuni esteri - Spagnolo - Tímalo ártico. Francese - Ombre arctique, Poisson bleu. Inglese - Arctic grayling, Back's grayling, Bluefish, Sailfin arctic grayling, American grayling, Siberian grayling, West Siberian grayling. Tedesco - Arktische Äsche. Olanda - Vlagzalm. Danimarca - Arktisk stalling, Stalling. Norvegia - Harr Norway. Svezia - Arktisk Harr. Finlandia - Harjus, Pohjanharjus. Polonia - Lipien syberyjski. Federazione Russa - Kharius sibirskiy, Sibirskii kharius. Lingua inuit (Eschimese) - Kewlook powak, Sulukpauga, Sulukpaugaq, Sulukpauvak, Sulupaugak, Tchulupa. Lingua Salish - K'wsech.

Thymallus arcticus Pallas, 1776. Reise durch verschiedene Provinzen des russischen Reiches. St. Petersburg. 3 vol. Sinonimo senior, nuova combinazione, nomenclatura ICZN valida.

Salmo (Truttac.) arcticus Pallas, 1776. Reise durch verschiedene Provinzen des russischen Reiches. St. Petersburg. 3 vol. Si vedano le osservazioni sulla sistematica in Kottelat 2006:23-24. Sinonimo senior, combinazione originale, nomenclatura ICZN non valida. Opinioni - Nomenclatura valida come Thymallus arcticus (Pallas 1776), vedere Berg 1948:423, Lee et al. 1980:120, McAllister 1990:83, Page & Burr 1991:47, Baxter & Stone 1995:200, Zhu 1995:12, Chereshnev 1996:603, Sigler & Sigler 1996:218, Kim 1997:355, Reshetnikov et al. 1997:732, Dorofeeva 1998:48, Scott & Crossman 1998:300, Fuller et al. 1999:266, Sheiko & Fedorov 2000:19, Lyons et al. 2000:67, Pietsch et al. 2001:143, Bogutskaya et al. 2001:46, Mecklenburg et al. 2002:191, Wydoski & Whitney 2003:103, Nelson et al. 2004:86, Bailey et al. 2004:159, Antonov 2004:445, Bogutskaya & Naseka 2004:147, Kottelat 2006:23, Ocock et al. 2006:50, Weiss et al. 2006:511, Scharpf 2006:34, Kottelat & Freyhof 2007:458, Knizhin & Weiss 2009:7. Località tipo - Affluenti del fiume Sob, bacino inferiore del fiume Ob vicino al 60°20' N. Tipo - Nessun esemplare noto.

Thymallus arcticus arcticus natio alchutovi Johansen, 1945. New forms of fishes of west Siberia. Tomskoe Otdelenie Moskovskago Obscestva Ispytatelej Priody. Tomsk N° 6: 1-16. Nome infrasubspecifico, non disponibile, nomenclatura ICZN non valida. Opinioni - In sinonimia di Thymallus arcticus (Pallas 1776), vedere Kottelat 2006:23. Provenienza del materiale - Lago Teletskoye, Siberia, Russia.

Salmo digitalis Bloch & Schneider, 1801. M. E. Blochii, Systema Ichthyologiae iconibus cx illustratum. Post obitum auctoris opus inchoatum absolvit, correxit, interpolavit Jo. Gottlob Schneider, Saxo. Berolini. Sumtibus Auctoris Impressum et Bibliopolio Sanderiano Commissum. M. E. Blochii, Systema Ichthyologiae.: i-lx + 1-584, Pls. 1-110. Nuovo nome non necessario per Salmo arcticus Pallas 1776, nomenclatura ICZN non valida. Opinioni - Sinonimo di Thymallus arcticus (Pallas 1776, vedere Kottelat 2006:23.

Thymallus arcticus lenensis Weiss, Knizhin, Kirillov & Froufe, 2006. Phenotypic and genetic differentiation of two major phylogeographical lineages of Arctic grayling Thymallus arcticus in the Lena River, and surrounding Arctic drainages. Biological Journal of the Linnean Society v. 88: 511-525. Nome non disponibile; proposto come nome "preliminare" per una popolazione di Thymallus del bacino del fiume Lena, Siberia, Russia. Località tipo - Bacino del fiume Lena, Russia. Tipo, non è stato stabilito nessun esemplare tipo.

Thymallus montanus Milner, 1874. Notes on the grayling of North America. Report of the United States Fish Commission Pt 2 (per il 1872-73): 729-742. Sinonimo junior, combinazione originale, nomenclatura ICZN non valida. Opinioni -Sinonimo di Thymallus arcticus (Pallas 1776), vedere Scott & Crossman 1998:304. Località tipo - Affluente del fiume Missouri a Camp Baker, Montana, U.S.A. Tipo - Nessuno. Sintipi - USNM 13090 (1), 13091 (1), 13092 (adesso 0).

Thymallus nikolskyi Kaschenko, 1899. Results of the Altai Zoological Expedition in 1899, Vertebrates. Pisces. Tomsk. Results of the Altai Zoological Expedition in 1899, Vertebrates. Pisces.: 131-155. Sinonimo junior, combinazione originale, nomenclatura ICZN non valida. Opinioni -Sinonimo di Thymallus arcticus (Pallas 1776), vedere Berg 1948:424, Dorofeeva 1998:48, Kottelat 2006:23, Knizhin & Weiss 2009:7. Località tipo - Fiume Tcharysh, fiume Katun (Lago Uimon), fiume Tom e corso superiore del fiume Kusnetzk negli Altai, Siberia, Russia. Tipo - Nessuno. Sintipi - (19) ubicazione ignota.

Thymallus nikolskyi var. ongudajensis Kaschenko, 1899. Results of the Altai Zoological Expedition in 1899, Vertebrates. Pisces. Tomsk. Results of the Altai Zoological Expedition in 1899, Vertebrates. Pisces.: 131-155. Sinonimo junior, combinazione originale, nomenclatura ICZN non valida. Opinioni -Sinonimo di Thymallus arcticus (Pallas 1776), vedere Berg 1948:424, Kottelat 2006:23. Località tipo - Fiume Urusul a Ongudai, affluente del fiume Katun, Altai, Siberia, Russia. Tipo - Nessuno. Sintipi - (8?) ubicazione ignota.

Thymalus ontariensis Valenciennes, 1848. In Cuvier & Valenciennes 1848. Histoire naturelle des poissons. Tome vingt et unième. Suite du livre vingt et unième et des Clupéoïdes. Livre vingt-deuxième. De la famille des Salmonoïdes. Histoire naturelle des poissons. v. 21: i-xiv + 1 p. + 1-536, Pls. 607-633. Catalogo - Bertin 1940:306. In origine il genere era probabilmente Thymallus. Sinonimo junior, combinazione originale, nomenclatura ICZN non valida. Opinioni -Sinonimo di Thymallus arcticus (Pallas 1776), vedere Baxter & Stone 1995:201, Scott & Crossman 1998:301. Se la località tipo fosse veramente il Lago di Ginevra l'attribuzione a T. arcticus sarebbe infondata (S. Porcellotti). Località tipo - "Lago Ontario" (probabilmente Lago di Ginevra, Svizzera). Tipo - nessuno. Sintipi - Da MNHN 0000-3662 a 0000-3663 (2).

Thymallus pallasii Valenciennes, 1848. In Cuvier & Valenciennes 1848. Histoire naturelle des poissons. Tome vingt et unième. Suite du livre vingt et unième et des Clupéoïdes. Livre vingt-deuxième. De la famille des Salmonoïdes. Histoire naturelle des poissons. v. 21: i-xiv + 1 p. + 1-536, Pls. 607-633. Catalogo - Bertin 1940:305. Sinonimo junior, combinazione originale, nomenclatura ICZN non valida. Opinioni -Sinonimo di Thymallus arcticus (Pallas 1776), ma valida sottospecie, vedere Berg 1948:428, Chereshnev 1996:603, Sigler & Sigler 1996:218, Reshetnikov et al. 1997:732, Dorofeeva 1998:48, Chereshnev et al. 2001:74 come pallasi, Fedorov et al. 2003:36, Weiss et al. 2006:511. Nomenclatura valida come Thymallus pallasii Valenciennes 1848, vedere Bogutskaya & Naseka 2004:148, Kottelat 2006:86. Località tipo - Fiume Kolyma, Russia. Tipo - Nessuno. Per le informazioni sul tipo vedere Svetovidov 1978:21. Sintipi - MNHN 0000-3664 (3), ZMB 23549 o 23550 (1).

Thymallus sellatus Kaschenko, 1899. Results of the Altai Zoological Expedition in 1899, Vertebrates. Pisces. Tomsk. Results of the Altai Zoological Expedition in 1899, Vertebrates. Pisces.: 131-155. Sinonimo junior, combinazione originale, nomenclatura ICZN non valida. Opinioni -Sinonimo di Thymallus arcticus (Pallas 1776), vedere Berg 1948:424, Kottelat 2006:23. Località tipo - Lago Tenga (Kenga), bacino del fiume Urusul nei monti Altai, Siberia, Russia. Tipo - Nessuno. Sintipi - (4) ubicazione ignota.

Coregonus signifer Richardson, 1823. Notice of the fishes. No. 6, in John Franklin. Narrative of a journey to the shores of the Polar Sea in 1819-22 ... with an appendix on various subjects relating to science and natural history. London. Narrative of a journey to the shores of the Polar Sea in 1819-22 ...: 705-728, Pls. 25-26. Sinonimo junior, combinazione originale, nomenclatura ICZN non valida. Opinioni - Sinonimo di Thymallus arcticus (Pallas 1776), ma valida sottospecie, vedere Chereshnev 1996:603, Reshetnikov et al. 1997:732, Dorofeeva 1998:48, Chereshnev et al. 2001:75. Sinonimo di Thymallus arcticus (Pallas 1776), vedere Baxter & Stone 1995:201, Scott & Crossman 1998:304, Mecklenburg et al. 2002:191. Menzioni - Kottelat 2006:86. Località tipo - Fiumi a nord del Gran Lago degli Schiavi, Canada. Tipo - Nessun esemplare noto.

Coregonus thymalloides Richardson, 1823. Notice of the fishes. No. 6, in John Franklin. Narrative of a journey to the shores of the Polar Sea in 1819-22 ... with an appendix on various subjects relating to science and natural history. London. Narrative of a journey to the shores of the Polar Sea in 1819-22 ...: 705-728, Pls. 25-26. Sinonimo junior, combinazione originale, nomenclatura ICZN non valida. Opinioni - Sinonimo di Thymallus arcticus (Pallas 1776), vedere Scott & Crossman 1998:304. Località tipo - Fiumi a nord del Gran Lago degli Schiavi, Territori del Nordovest, Canada. Tipo - Nessun esemplare noto.

Thymallus tricolor Cope, 1865. Partial catalogue of the cold-blooded Vertebrata of Michigan. Part II. Proceedings of the Academy of Natural Sciences of Philadelphia v. 17: 78-88. Catalogo - Böhlke 1984:150; vedere anche Fowler 1918:5. Sinonimo junior, combinazione originale, nomenclatura ICZN non valida. Opinioni - Sinonimo di Thymallus arcticus (Pallas 1776), vedere Baxter & Stone 1995:201, Scott & Crossman 1998:304. Località tipo - Fiume Au Sable, Michigan, U.S.A. Tipo - Olotipo ANSP 7796. Possibili esemplari tipo tra gli specimen di Cope - ?UMMZ 157347 (1), 213814 (1).

Thymallus articus yaluensis Mori, 1928. On the fresh water fishes from the Yalu River, Korea, with descriptions of new species. Journal of the Chosen Natural History Society No. 6: 54-70 + map. Scritto jaluensis nel titolo principale a p. 57 e yaluensis a p. 22; nome tratto da quello del fiume Yalu, così yaluensis è considerato scritto correttamente. Opinioni - Sinonimo di Thymallus arcticus (Pallas 1776), vedere Berg 1948:430. Sinonimo di Thymallus arcticus (Pallas 1776), ma valida sottospecie, vedere Zhu 1995:12, Kim 1997:355 (scritto jaluensis), Kim & Park 2002:275 (scritto jaluensis), Youn 2002:194, 531 (scritto jaluensis). Località tipo - Corso superiore del fiume Yalu a Kozan, Corea. Tipo - Un olotipo ma con ubicazione ignota. Esemplari addizionali - (10).

Fonti bibliografiche principali - Kottelat M. and J. Freyhof 2007. Handbook of European freshwater fishes. Kottelat, Cornol, Switzerland and Freyof, Berlin, Germany. Pagina 458 - 459.

Bibliografia - Andriyashev, A. P. & N. V. Chernova 1995. Annotated list of fishlike vertebrates and fish of the arctic seas and adjacent waters.

Andriyashev, A. P. 1964. Fishes of the northern seas of the U.S.S.R.

Anonymous 1996. Fish collection database of the University of British Columbia Fish Museum.

Anonymous 1999. Fish collection database of the Natural History Museum, London (formerly British Museum of Natural History (BMNH)).

Anonymous 2000. Fish collection database of the J. L. B. Smith Institute of Ichthyology, Grahamstown, South Africa.

Anonymous 2001. Fish collection database of the National Museum of Natural History (Smithsonian Institution).

Anonymous 2001. Fish collection database of the Zoological Museum, University of Copenhagen.

Anonymous 2002. Fish collection database of the American Museum of Natural History.

Anonymous 2002. Fish collection of the Royal British Columbia Museum.

Anonymous 2003. Fish collection of the Royal Ontario Museum.

Arkhipchuk, V. V. 1999. Chromosome database.

Berg, L. S. 1936. Note on Coregonus (Prosopium) cylindraceus (Pallas).

Berg, L. S. 1962. Freshwater fishes of the U.S.S.R. and adjacent countries. volume 1, 4th edition.

Bishop, F. G. 1971. Observations on spawning habits and fecundity of the Arctic grayling.

Blanc, M., J.-L. Gaudet, P. Banarescu & J.-C. Hureau 1971. European inland water fish. A multilingual catalogue.

Bogutskaya, N. G. & A. M. Naseka 2004. Catalogue of agnathans and fishes of fresh and brackish waters of Russia with comments on nomenclature and taxonomy.

Breder, C. M. and D. E. Rosen 1966. Modes of reproduction in fishes.

Brown, C. J. D. 1938. Observations on the life-history and breeding habits of the Montana grayling.

Byorth, P. A. & J. P. Magee 1998. Competitive interactions between Arctic grayling and brook trout in the Big Hole River drainage, Montana.

Carl, G. C., W. A. Clemens & C. C. Lindsey 1959. The freshwater fishes of British Columbia (3rd revision).

Carl, H. 2003. Danish fish names.

Chinese Academy of Fishery Sciences 2003. Chinese aquatic germplasm resources database.

Clark, B., D. M. Flemming & W. P. Ridder 1991. Stock assessment of Arctic grayling in the Salcha, Chatanika, and Goodpaster rivers.

Clark, R. A. 1990. Stock status of Chena River Arctic grayling during 1990.

Clark, R. A. 1993. Stock assessment of Arctic grayling in Fielding Lake during 1991 and 1992.

Coker, G. A., C. B. Portt & C. K. Minns 2001. Morphological and ecological characteristics of Canadian freshwater fishes.

Cope, E. D. 1865. Partial catalogue of the cold-blooded vertebrata of Michigan. Part 2.

Craig, P. C. 1984. Fish use of coastal waters of the Alaskan Beaufort Sea: a review.

Dall, W. H. 1870. Alaska and its resources.

Danko, J. P. 1998. Building a reliable database from a native oral tradition using fish-related terms from the Saanich language.

DeCicco, A. L., M. F. Merritt & A. E. Bingham 1997. Characteristics of a lightly exploited population of Arctic grayling in the Sinuk River, Seward Peninsula, Alaska.

Dulmaa, A. 1999. Fish and fisheries in Mongolia.

Evermann, B. W. & E. L. Goldsborough 1907. The fishes of Alaska.

FAO-FIES 2008, Aquatic Sciences and Fisheries Information System (ASFIS) species list.

FAO-FIES 2010. Aquatic Sciences and Fisheries Information System (ASFIS) species list.

Fedorov, V. V., I. A. Chereshnev, M. V. Nazarkin, A. V. Shestakov & V. V. Volobuev 2003. Catalog of marine and freswater fishes of the northern part of the Sea of Okhotsk.

Fisheries Branch 1994. Freshwater Fishing Regulations Synopsis, 1994-1996.

Fleming, D. F. 1991. Stock assessment of Arctic grayling in Piledriver Slough, 1991.

Freyhof, J. & Kottelat, M. 2008. Thymallus arcticus. In: IUCN 2010. IUCN Red List of Threatened Species.

Frimodt, C. 1995. Multilingual illustrated guide to the world's commercial coldwater fish.

Grabda, E. & T. Heese 1991. Polskie nazewnictwo popularne kraglouste i ryby. Cyclostomata et Pisces.

Graves, J. & E. Hall 1988. Arctic Animals.

Günther, A. 1866. Catalogue of the fishes of the British Museum, Vol. VI.

Hemming, C. R. 1997. Experimental introduction of Arctic grayling to a rehabilitated gravel extraction site, North Slope, Alaska.

Hubbs, C. L. & K. F. Lagler 1947. Fishes of the Great Lakes Region.

Hugg, D. O. 1996. MAPFISH georeferenced mapping database. Freshwater and estuarine fishes of North America.

IGFA 2001, Database of IGFA angling records until 2001.

Institute of Hydrobiology, Academia Sinica, Shanghai Natural Museum and Ministry of Agriculture of China 1993. The freshwater fishes of China in coloured illustrations. Vol. 3.

International Game Fish Association 1991. World record game fishes.

IUCN. 2008. 2008 IUCN Red List of Threatened Species. http://www.iucnredlist.org.

Jang, M.-H., J.-G. Kim, S.-B. Park, K.-S. Jeong, G.-I. Cho & G.-J. Joo 2002. The current status of the distribution of introduced fish in large river systems of South Korea.

Jelks, H. L., S. J. Walsh, N. M. Burkhead, S. Contreras-Balderas, E. Díaz-Pardo, D. A. Hendrickson, J. Lyons, N. E. Mandrak, F. McCormick, J. S. Nelson, S. P. Platania, B. A. Porter, C. B. Renaud, J. J. Schmitter-Soto, E. B. Taylor & M. L. Warren, Jr. 2008. Conservation status of imperiled North American freshwater and diadromous fishes.

Jenness, D. 1928. Eskimo language and technology.

Klinkhardt, M, M. Tesche & H. Greven 1995. Database of fish chromosomes.

Koskinen, M. T., Knizhin, I., Primmer, C. R., Schlötterer, C. & Weiss, S. 2002. Mitochondrial and nuclear DNA phylogeography of Thymallus spp. (grayling) provides evidence of ice-age mediated environmental perturbations in the world's oldest body of freshwater, Lake Baikal.

Kottelat, M. & J. Freyhof 2007. Handbook of European freshwater fishes.

Kottelat, M. 2006. Fishes of Mongolia. A check-list of the fishes known to occur in Mongolia with comments on systematics and nomenclature.

Kruse, T. E. 1959. Grayling of Grebe Lake, Yellowstone National Park, Wyoming.

Laird, J. A. 1928. Grayling in the east.

Levanidov, V. Y. 1959. Feeding and trophic relations of fishes in foothill inflows of lower Amur.

Lord, R. F. 1932. Notes on Montana grayling at the Pittsford, Vt., experimental trout hatchery.

Machacek, H. (ed.) 2006. World Records Freshwater Fishing.

Machacek, H. (ed.) 2009. World Records Freshwater Fishing.

McAllister, D. E. 1990. A list of the fishes of Canada.

McAllister, D. E., V. Legendre & J. G. Hunter 1987. Liste de noms inuktitut (esquimaux), français, anglais et scientifiques des poissons marins du Canada arctique.

McClanes, A. J. (ed.) 1974. Field guide to freshwater fishes of North America.

McPhail, J. D. & R. Carveth 1993. Field key to the freshwater fishes of British Columbia.

McPhail, J. D. and C. C. Lindsey 1970. Freshwater fishes of northwestern Canada and Alaska.

Milner, J. W. 1874. Notes on the grayling on North America.

Moring, J. R. 1996. Fish discoveries by the Lewis and Clark and Red River Expeditions.

Morrow, J. E. 1980. The freshwater fishes of Alaska.

Murdoch, J. 1892. Ethnological results of the Point Barrow expedition.

National Park Service, U.S. Department of the Interior 2008. Freshwater fishes of Wrangell-St. Elias National Park and Preserve, Alaska.

Nelson, J. S., E. J. Crossman, H. Espinosa-Pérez, L. T. Findley, C. R. Gilbert, R. N. Lea & J. D. Williams 2004. Common and scientific names of fishes from the United States, Canada, and Mexico.

Nelson, P. H. 1954. Life history and management of the American grayling (Thymallus signifer tricolor) in Montana.

Nikolsky, G. W. 1957. Spezielle Fischkunde.

Norden, G. R. 1959. Comparative morphology of certain salmonid fishes with particular reference to the grayling (Thymallus arcticus), and its phylogeny.

Organisation for Economic Co-operation and Development 1990. Multilingual dictionary of fish and fish products.

Page, L. M. & B. M. Burr 1991. A field guide to freshwater fishes of North America north of Mexico.

Pallas, P. S. 1776. Reise durch verschiedene Provinzen des Russischen Reichs (1768-74).

Pauly, D. 1978. A preliminary compilation of fish length growth parameters.

Petitot, E. 1876. Vocabulaire français-esquimau.

Pietsch, T. W., K. Amaoka, D. E. Stevenson, E. L. MacDonald, B. K. Urbain & J. A. López 2000. Freshwater fishes of the Kuril Islands and adjacent regions.

Pivnicka, K. & Hensel, K. 1978. Morphological variation in the genus Thymallus Cuvier, 1829 and recognition of the species and subspecies.

Podlesnyi, A. V. 1958. Fishes of Enisey, their environments and use.

Quast, J. C. & E. L. Hall 1972. List of fishes of Alaska and adjacent waters with a guide to some of their literature.

Randolph, S. & M. Snyder 1993. The seafood list: FDA's guide to acceptable market names for seafood sold in interstate commerce.

Rass, T. S. 1983. Fish.

Reed, R. J. 1964. Life history and migration patterns of Arctic grayling, Thymallus arcticus (Pallas), in the Tanana River drainage of Alaska.

Rempel, L. L. & D. G. Smith 1998. Postglacial fish dispersal from the Mississippi refuge to the Mackenzie River basin.

Reshetnikov, Y. S., N. G. Bogutskaya, E. D. Vasil'eva, E. A. Dorofeeva, A. M. Naseka, O. A. Popova, K. A. Savvaitova, V. G. Sideleva & L. I. Sokolov 1997. An annotated check-list of the freshwater fishes of Russia.

Richardson, J. 1836. The fish.

Robins, C. R., R. M. Bailey, C. E. Bond, J. R. Brooker, E. A. Lachner, R. N. Lea & W. B. Scott 1980. A list of common and scientific names of fishes from the United States and Canada.

Robins, C. R., R. M. Bailey, C. E. Bond, J. R. Brooker, E. A. Lachner, R. N. Lea & W. B. Scott 1991. Common and scientific names of fishes from the United States and Canada.

Schallock, E. W. 1966. Grayling life history related to a hydroelectric development on the Chatanika River, Alaska.

Scott, W. B. & E. J. Crossman 1973. Freshwater fishes of Canada.

Scott, W. B. 1958. A checklist of the freshwater fishes of Canada and Alaska.

Severin, S. O. & E. A. Zinov'yev 1982. Karyotypes of isolated populations of Thymallus arcticus from the Ob' River basin.

Stolbov, A. Y. & Y.S. Alikin 1977. Temperature dependence of active metabolism and swimming speed of the Baikal grayling, Thymallus arcticus baicalensis.

Swedish Museum of Natural History 1999. NRM Ichthyology collection database.

Thurston, R. V. & P. C. Gehrke 1993. Respiratory oxygen requirements of fishes: description of OXYREF, a data file based on test results reported in the published literature.

Vascotto, G. L. 1970. Summer ecology and behavior of the grayling of McManus Creek, Alaska.

Vasil'ev, V.P. 1980 Chromosome numbers in fish-like vertebrates and fish.

Walker, K. F. & H. Z. Yang 1999. Fish and fisheries in western China.

Walters, V. 1955. Fishes of the western arctic America and eastern arctic Siberia, taxonomy and zoogeography.

Ward, J. C. 1951. The breeding biology of the Arctic grayling in the southern Athabasca Drainage.

Webster, D. H. & W. Zibell 1970. Inupiat Eskimo dictionary.

Wheeler, A. 1977. Das grosse Buch der Fische.

Whitehouse, F. C. 1919. Notes on some fishes of Alberta and adjacent waters.

Wojcik, F. J. 1955. Life history and management of the grayling in interior Alaska.

Wynne-Edwards, V. C. 1952. Fishes of the Arctic and subarctic.

Zinoviev, E. A. 1976. Parallelisms in variation of European and Siberian graylings. Zoolological Institute, Academy of Sciences of USSR, Leningrad.

Zinoviev, E.A. 1989. Hybridization of graylings in the zone of sympatry. Academy of Sciences of USSR, Sverdlovsk.